|Photo by Jim Craig|
The presence of the vertebrae of aquatic snakes in the Ak-Kaya locality was first mentioned by Bratishko and Udovichenko in 2007. Later, one vertebra of Palaeophis sp. was described from this locality by Snetkov and Bannikov 2010. In 2010, E.A. Zvonok made an exploratory excavation pit about 500 m to the east from the nummulite limestone quarry of Ak-Kaya and collected six vertebrae of Palaeophis nessovi. In 2011, E.A. Zvonok excavated the second pit and recovered numerous vertebrae of aquatic snakes. This material is discussed in a new paper by Zvonok and Snetkov (2012).
They refer the vertebrae to three known species of the genus Palaeophis (P. tamdy, P. nessovi and P. cf. toliapicus). These species demonstrate different degree of aquatic adaptations and lived in a restricted area in the eastern part of the Tethys Ocean during the Middle Eocene.
Palaeophis nessovi has well developed pterapophyses, comparatively short prezygapophyses and strongly laterally compressed posterior and anterior trunk vertebrae. These characters suggest that P. nessovi was well adapted to the aquatic mode of life . Palaeophis tamdy has large prezygapophyses and lacks pterapophyses, and is considered less specialized. Palaeophis cf. toliapicus, was most likely adapted to an aquatic life style even better than P. nessovi according to less developed prezygapophyses and stronger lateral compression of the vertebra.
Zvonok E.A. and P.B. 2012. Snetkov NEW FINDINGS OF SNAKES OF THE GENUS PALAEOPHIS OWEN, 1841 (ACROCHORDOIDEA: PALAEOPHIIDAE) FROM THE MIDDLE EOCENE OF CRIMEA. Proceedings of the Zoological Institute RAS 316:392-400.