Phytolophis punctata
Gray,1849 Catalogue of the Specimens of Snakes in the Collection of the British
Museum, p. 68. Type locality "India" (in error). Holotype: BMNH 1946.1.2.37.
Eurostus heteraspis
Bleeker, 1859 Natuurkundig Tijdshrift voor Nederlandsch Indië, 14:440. Type
locality: Sinkawang, Borneo. Collector: Dr. Bleeker. Holotype: BMNH
1946.1.2.39.
Tachyplotus hedemanni
Reinhardt, 1866 Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i
Kjobenhaun, p. 151. Type locality: Billiton Is. [Indonesia]. Holotype: ?RNHM
Pythonopsis borneensis
Peters, 1871 Monatsberichte der Preussischen Akademie der Wissenschaften zu
Berlin, p. 576. Type locality: Sarawak. Holotype: Berlin Museum?
Hypsirhina hageni
Lidth de Jeude, 1890 Notes from the Leyden Museum, 12:20. Pl. 1. Type locality:
“Laboean.” David and Vogel (1996:118) note that the type locality given by
Lidth de Jeude for it as Laboean or Lubuan is now known as Belawan, Sumatra
[3°47’N 98°41’E]. This city is on the northeast coast of Sumatera Utara. Holotype: RMNH 862.
Hypsirhina punctata
- Boulenger, 1896 Catalogue of Snakes in the British Museum 3:12.
Enhydris punctata
- Kinghorn, 1929 Snakes of Australia, p. 89. [probably based upon a
misidentified Fordonia.]
This is in all likelihood a collection of several species
that are isolated in peat swamp forests and associated streams across the range
of the species group. It seems probable that one or more of the names in the
synonymy above are valid, but because specimens are relatively rare and many
specimens lack specific locality data it is not possible to currently sort
these out.
Etymology: The name punctata may have originated from the
Latin punctum, meaning “spot,” however this snake does not have a spotted
pattern, but it does have yellow cross bars. It seems more likely it is derived
from the Latin punctura, which means “hole” or “prick”, or “puncture” and
refers to its bite. Common Names: The Phytolopsis (Gray, 1849); Spotted
Water-snake (Kinghorn, 1929).
Distribution: This is a species endemic to the Sunda Shelf
and peninsular Malaysia, it may eventually be found in extreme southern
Thailand. Thus, it ranges from Sumatra and the west coast of peninsular
Malaysia eastward to Borneo. Confusion about this species’ presence in
Australia is widespread in the literature. Kinghorn (1929) appears to be the
first person to suggest its presence in Australia, he wrote, "During
recent years this species has been found in north Australia but previously it
was thought to be restricted to Sumatra, Borneo and neighbouring islands."
Cogger (1975) had a map for it in his first edition of Reptiles and Amphibians
of Australia, but omitted it from latter editions. Gow (1977, plate 7) has a
photo labeled Enhydris punctata from
Darwin, Australia. It is in fact a specimen of Fordonia.
Size: The largest specimen measured was 760 mm in total
length with a 95 mm tail. David and Vogel (1996) wrote: “This small [snake] has
a maximum recorded length of 450 mm, with adults averaging 340 - 400 mm.” Similarly,
Ernst and Zug (1996) have a table in Appendix 2 that gives the maximum length
of this species as 0.45 m. However, adults of these snakes are massively built
and reach at least a length 760 m, but most of the museum specimens available
are juveniles. Tails on males are 19 - 21% of the SVL, while females have tails
that are 8.5 - 17.8% of the SVL.
External Morphology: The head is slightly distinct from
neck; this seems to be true for smaller specimens rather than larger ones. The
overall body shape is cylindrical and slightly depressed anteriorly, including
the head. The eyes are small; the diameter is about 1.5 times the eye-nostril
distance. The eyes are more dorsal than dorsolateral in position. On the head the
rostral scale is pentagonal, barely visible from above, and about as broad and
it is tall. The nasals are semi-divided with the cleft touching the first
labial. The nasals scales have a diameter that is equal to the eye diameter;
these and most of the other head scales are tuberculate. The internasal is
small and partially penetrates the seam between the nasals; it is about one
fourth of a nasal scale in area; in one specimen from Borneo (MCZ 5165) it is
divided. It is quadrangular. Each of the prefrontals is about twice the size of
the internasal and each is in contact with a loreal. The frontal’s length is
about equal to the interorbital distance. It is triangular with the apex
pointing posteriorly. The parietals are about equal in length to the frontal.
These and most of the other head scales are imbricate. The loreal may be single
or divided (it is single in the type specimen), with the anterior section being
twice as broad as the posterior section. It is in contact with upper labials 1
- 4 or 1 - 5. The single supraocular is narrower on the front margin than it is
on the posterior margin. The preocular can be single or divided. There are two
postocular scales, with the ventral scale extending slightly under the orbit to
form a partial subocular. There are no suboculars. The temporal formula is 1 +
2; the tertiary temporal scales are indistinguishable from occipitals. There
are 12 - 14 upper labials; the first five or six are anterior to the orbit,
overlap each other and are very narrow at the point the contact the loreal. The
sixth or seventh is wider, enters the orbit and is divided into two tiers. The
posterior labials are divided so that they form three or four tiers of scales.
At the 11th labial the angle of the lower jaw turns upward. On the chin the
lower labials number 12 - 15. The first pair forms the mental groove in most
specimens, but in the type specimen (BM 1946.1.2.37) the first two lower
labials form the mental groove. The first 5 are in contact with the anterior
pair of chin shields. The sixth is the largest. The anterior pair of chin
shields are described by Gyi as “petal shaped” – meaning that they are very
broad, almost as broad as they are long. The second pair is greatly reduced and
separated by a pair of smaller scales. The gulars number seven. On the body the
dorsal scales are in 23 - 27 rows on the neck, at mid-body they number 25 - 27,
and near the vent the rows are reduced to 21 - 23 rows. The first several rows
are ovate and they become slightly more elongated toward the midline; they are
smooth and iridescent. Smedley (1931) reported five specimens (possibly
immature) in the Raffle’s Museum collected on the east coast of the east coast
of the Malay Peninsula, all of which have 23 scale rows, I have not seen any
specimens with 23 rows at mid-body. The ventrals are broad, about 3 - 3.5 times
the height of a nearby dorsal scale. They number 148 - 160 (148 - 151 in males,
155 - 160 in females).The anal plate is divided and slightly longer than the
preceding ventral. On the tail dorsal scales on the tail are ovate. The
subcaudal scales are divided and number 27 - 46 (32 - 44 in females, 46 - 48 in
males). Three males have tails that are 0.19 - 0.20 of the SVL; nine females
have tails that are 0.09 - 0.21 of the SVL. At the base of the tail the width
is 94% of the height based upon the average of five specimens. The crown of the
head is black with a yellow interorbital stripe that is indistinct. The dorsum
is black with the exception of a faint occipital stripe and incomplete
transverse bars on the anterior of the body, which may appear as yellow spots.
Scale rows one and two are mostly yellow with some invasion of dark pigment.
Scale rows 3 - 5 are black with yellow spots or solid black. The lateral areas
of the head have some yellow blotches and the snout is black; the chin and
ventral are yellow; the ventral tail is yellow with a mid-black line.
Natural History: Museum notes for peninsular Malaysian
specimens (FMNH250111-2) state that they were collected in “blackwater
streams.” And a specimen collected at Pondak Tanjung Forest Reserve, Selama
Perak, Malaysia on 1 March 2001 by Shahrul Anuaroast came from a peat swamp
forest. These notes suggest that this uncommon snake is most likely a peat
swamp or blackwater habitat specialist. The fragmented nature of this habitat
and the high level of fish endemism in the peat swamps of Southeast Asia add an
interesting dimension to this species complex, and to homalopsid diversity. The
details of its life style remains virtually unknown. One specimen, MCZ 5165,
contained 15 yolked follicles.